Blepharisma

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Blepharisma is a genus of unicellular ciliate protists found in fresh and salt water. The group includes about 40 accepted species, and many sub-varieties and strains. While species vary considerably in size and shape, most are easily identified by their red or pinkish color, which is caused by granules of the pigment blepharismin. • Like all ciliates, Blepharisma reproduce asexually, by binary fission, dividing transversally. Fission may occur spontaneously, as part of the vegetative cell cycle, or it may follow a sexual phenomenon called conjugation, a process through which genetic material is exchanged between cells. In conjugation, two organisms come into close contact, and a temporary cytoplasmic bridge forms between them. The micronuclei of each cell then undergo meiosis, and haploid micronuclei pass from one individual to the other. This permits the reshuffling of hereditary characteristics, as in other types of sexual reproduction. Conjugation is immediately followed by binary fission of the two conjugants. • Blepharisma are markedly photophobic, and when light levels are increased will seek out darkened areas. The ability to detect light is accomplished with photosensitive pigment granules located just under the plasma membrane of the cell. The pigment in these granules is blepharismin, the same substance that gives Blepharisma their characteristic pinkish color. Blepharisma are usually pink when collected in nature, but when grown in darkness with abundant food they turn red. Exposure to light or starvation causes them to lose their color, but deeply-pigmented cells can even be killed by strong 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