DNA Replication Model

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Transcript: • This is a model of DNA, a double strand of DNA, and when it's replicated, it is replicated semi-conservatively. So that means that these two strands are the parental strands, or the templates. And it gets split, and then they actually build up two new strands off of that that are complimentary. So, these are the base pairs and you have guanine that pairs with cytosine, and adenine that pairs with cytosine. And those are the only two combinations you can have, you can't have a yellow and a blue because if you look at their shapes, they actually won't fit together. That's just how they work. • Camera change* • So the reason that the nucleotides can only pair in these ways is because you have purines and pyrimidines. The purines are adenine and guanine, and they have two carbon rings. And the pyrimidines are cytosine and thymine and they have a single carbon ring. When you bond them together, you'll take and adenine, which is the red piece, and that will pair with thymine, which is the yellow piece. So when we connect them you're having one purine bond with one pyrimidine. And these two actually form- they would actually go this way, but that was a mistake. So when you have these two together, they have 3 hydrogen bonds (THEY REALLY HAVE TWO) between them. And when you bond the thymine (I misspoke, it is cytosine) and the guanine, they bond like this and they have two hydrogen bonds (THEY REALLY HAVE THREE) between them, and that's how they pair together. • Camera change* • So this is the double stranded DNA after it has been split open by helicase. So the helicase come in like this and it creates a replication fork. So DNA is always built 5’ to 3’, so here is the 5’ end for this strand, and it goes 5’ to 3’. Here's the 5’ end here, and it goes 5’ to 3’. So when you are building the compliments, you have to build them the same type of way. • So here, this is called the leading strand. And the leading strand is easy, because right here it's going 5 to 3, so the compliment is going to go 5 to 3 like this. So what happens is, DNA polymerase comes on and it attaches the pieces. It finds the matching nucleotide bases and it adds them on into a complimentary strand just like this. That is DNA polymerase. • Camera change* • So this is what the leading strand looks like when you add most of the nucleotides on. The next strand, this is called the lagging strand, and it is a little more difficult because here we have it going 5’ to 3’ on the template. But the complementary strand has to go 5’ to 3’ like this. So, since the helicase hasn't finished unzipping the whole thing, it can't start over here because this isn't undone yet. So they're going to be built in fragments which are called Okazaki fragments. So the DNA polymerase comes in, and starts attaching nucleotides on, just working its way outwards (from the fork) because it has to build 5’ to 3’. And then what happens when it gets to the end, after, the DNA helicase is going to keep unzipping this original template, so it's gonna move out. The leading strand is just going to keep on building forward this way, the lagging strand is just going to come out and it's going to start here and work its way back towards its beginning. When you get here, we’re not going to fill in the whole thing, but, say I filled in all of these and I got to here, this model shows that they're connected, but what really happens is there's a little bit of a gap. So DNA ligase will come in, it's going to repair this hole here, and it's going to make it so that all these fragments are connected so when you get to the end… • Text screen • Camera change* • You have these two different strands. So if you look, this one here and the outside one here were originally attached but we split them through the replication fork, and we made the leading strand here and Okazaki fragments going this way on the lagging strand. • Real DNA is thousands of base pairs long, and these ones are only 22 so it would actually take a little bit longer than this took, but after you're done then DNA replication is over, and it's called semi- conservative remember because we have these two which were from the original so they were conserved, and then you're creating two new strands that are the complementary strands with new nucleotides added on. • 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